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Network neighborhood for query Sequence : VMD2_HUMAN

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VMD2_HUMAN
SwissProt Accessions	O76090, O75904, Q8IUR9, Q8IZ80	Gene names	VMD2
Domain Architecture
Description	Bestrophin-1 (Vitelliform macular dystrophy protein 2) (TU15B).
Rank Plots	Jump to hits sorted by θ Jump to hits sorted by NC score

Hits sorted by θ

VMD2_HUMAN
θ value	0 (rank : 1)	NC score	0.999962 (rank : 1)
Query Neighborhood Hits	26	Target Neighborhood Hits	26	Shared Neighborhood Hits	26
SwissProt Accessions	O76090, O75904, Q8IUR9, Q8IZ80	Gene names	VMD2
Domain Architecture
Description	Bestrophin-1 (Vitelliform macular dystrophy protein 2) (TU15B).

VMD2_MOUSE
θ value	0 (rank : 2)	NC score	0.992437 (rank : 2)
Query Neighborhood Hits	26	Target Neighborhood Hits	15	Shared Neighborhood Hits	7
SwissProt Accessions	O88870, Q6H1V0	Gene names	Vmd2, Bmd1
Domain Architecture	No domain graphic available
Description	Bestrophin-1 (Vitelliform macular dystrophy protein 2 homolog).

VMDL3_HUMAN
θ value	3.50381e-145 (rank : 3)	NC score	0.977047 (rank : 7)
Query Neighborhood Hits	26	Target Neighborhood Hits	32	Shared Neighborhood Hits	7
SwissProt Accessions	Q8N1M1, Q8N356, Q8NFT9, Q9BR80	Gene names	VMD2L3
Domain Architecture
Description	Bestrophin-4 (Vitelliform macular dystrophy 2-like protein 3).

VMDL3_MOUSE
θ value	3.50381e-145 (rank : 4)	NC score	0.978253 (rank : 6)
Query Neighborhood Hits	26	Target Neighborhood Hits	30	Shared Neighborhood Hits	8
SwissProt Accessions	Q6H1V1	Gene names	Vmd2l3
Domain Architecture	No domain graphic available
Description	Bestrophin-4 (Vitelliform macular dystrophy 2-like protein 3).

VMDL1_HUMAN
θ value	2.35023e-141 (rank : 5)	NC score	0.983462 (rank : 5)
Query Neighborhood Hits	26	Target Neighborhood Hits	19	Shared Neighborhood Hits	7
SwissProt Accessions	Q8NFU1, Q9NXP0	Gene names	VMD2L1
Domain Architecture
Description	Bestrophin-2 (Vitelliform macular dystrophy 2-like protein 1).

VMDL1_MOUSE
θ value	2.35023e-141 (rank : 6)	NC score	0.984085 (rank : 4)
Query Neighborhood Hits	26	Target Neighborhood Hits	19	Shared Neighborhood Hits	7
SwissProt Accessions	Q8BGM5, Q8VCM0	Gene names	Vmd2l1
Domain Architecture
Description	Bestrophin-2 (Vitelliform macular dystrophy 2-like protein 1).

VMDL2_HUMAN
θ value	2.69525e-129 (rank : 7)	NC score	0.985886 (rank : 3)
Query Neighborhood Hits	26	Target Neighborhood Hits	8	Shared Neighborhood Hits	7
SwissProt Accessions	Q8NFU0	Gene names	VMD2L2
Domain Architecture
Description	Bestrophin-3 (Vitelliform macular dystrophy 2-like protein 2).

ATE1_HUMAN
θ value	0.0961366 (rank : 8)	NC score	0.046737 (rank : 8)
Query Neighborhood Hits	26	Target Neighborhood Hits	11	Shared Neighborhood Hits	2
SwissProt Accessions	O95260, O95261, Q8WW04	Gene names	ATE1
Domain Architecture
Description	Arginyl-tRNA--protein transferase 1 (EC 2.3.2.8) (R-transferase 1) (Arginyltransferase 1) (Arginine-tRNA--protein transferase 1).

CYTSA_HUMAN
θ value	0.0961366 (rank : 9)	NC score	0.017808 (rank : 11)
Query Neighborhood Hits	26	Target Neighborhood Hits	1138	Shared Neighborhood Hits	5
SwissProt Accessions	Q69YQ0, O15081	Gene names	CYTSA, KIAA0376
Domain Architecture	No domain graphic available
Description	Cytospin-A (NY-REN-22 antigen).

UBP10_HUMAN
θ value	0.163984 (rank : 10)	NC score	0.026889 (rank : 9)
Query Neighborhood Hits	26	Target Neighborhood Hits	153	Shared Neighborhood Hits	3
SwissProt Accessions	Q14694, Q9BWG7, Q9NSL7	Gene names	USP10, KIAA0190
Domain Architecture
Description	Ubiquitin carboxyl-terminal hydrolase 10 (EC 3.1.2.15) (Ubiquitin thioesterase 10) (Ubiquitin-specific-processing protease 10) (Deubiquitinating enzyme 10).

SPEG_HUMAN
θ value	0.365318 (rank : 11)	NC score	0.004118 (rank : 24)
Query Neighborhood Hits	26	Target Neighborhood Hits	1762	Shared Neighborhood Hits	7
SwissProt Accessions	Q15772, Q27J74, Q695L1, Q6FGA6, Q6ZQW1, Q6ZTL8, Q9P2P9	Gene names	SPEG, APEG1, KIAA1297
Domain Architecture
Description	Striated muscle preferentially expressed protein kinase (EC 2.7.11.1) (Aortic preferentially expressed protein 1) (APEG-1).

PLXB1_MOUSE
θ value	0.47712 (rank : 12)	NC score	0.013462 (rank : 15)
Query Neighborhood Hits	26	Target Neighborhood Hits	259	Shared Neighborhood Hits	3
SwissProt Accessions	Q8CJH3, Q6ZQC3, Q80ZZ1	Gene names	Plxnb1, Kiaa0407
Domain Architecture	No domain graphic available
Description	Plexin-B1 precursor.

CYTSA_MOUSE
θ value	0.62314 (rank : 13)	NC score	0.015434 (rank : 13)
Query Neighborhood Hits	26	Target Neighborhood Hits	1104	Shared Neighborhood Hits	3
SwissProt Accessions	Q2KN98, Q8CHF9	Gene names	Cytsa, Kiaa0376
Domain Architecture	No domain graphic available
Description	Cytospin-A.

TULP4_MOUSE
θ value	1.06291 (rank : 14)	NC score	0.020710 (rank : 10)
Query Neighborhood Hits	26	Target Neighborhood Hits	247	Shared Neighborhood Hits	4
SwissProt Accessions	Q9JIL5, Q8CA75, Q922C2	Gene names	Tulp4, Tusp
Domain Architecture
Description	Tubby-like protein 4 (Tubby superfamily protein).

IF4G1_HUMAN
θ value	1.38821 (rank : 15)	NC score	0.015539 (rank : 12)
Query Neighborhood Hits	26	Target Neighborhood Hits	347	Shared Neighborhood Hits	5
SwissProt Accessions	Q04637, O43177, O95066, Q5HYG0, Q6ZN21, Q8N102	Gene names	EIF4G1, EIF4G, EIF4GI
Domain Architecture
Description	Eukaryotic translation initiation factor 4 gamma 1 (eIF-4-gamma 1) (eIF-4G1) (eIF-4G 1) (p220).

CSPG2_MOUSE
θ value	2.36792 (rank : 16)	NC score	0.007615 (rank : 20)
Query Neighborhood Hits	26	Target Neighborhood Hits	711	Shared Neighborhood Hits	5
SwissProt Accessions	Q62059, Q62058, Q9CUU0	Gene names	Cspg2
Domain Architecture
Description	Versican core protein precursor (Large fibroblast proteoglycan) (Chondroitin sulfate proteoglycan core protein 2) (PG-M).

EFS_MOUSE
θ value	3.0926 (rank : 17)	NC score	0.014075 (rank : 14)
Query Neighborhood Hits	26	Target Neighborhood Hits	389	Shared Neighborhood Hits	8
SwissProt Accessions	Q64355	Gene names	Efs, Sin
Domain Architecture
Description	Embryonal Fyn-associated substrate (SRC-interacting protein) (Signal- integrating protein).

NCOR2_MOUSE
θ value	3.0926 (rank : 18)	NC score	0.010198 (rank : 19)
Query Neighborhood Hits	26	Target Neighborhood Hits	790	Shared Neighborhood Hits	4
SwissProt Accessions	Q9WU42, Q9WU43, Q9WUC1	Gene names	Ncor2, Smrt
Domain Architecture
Description	Nuclear receptor corepressor 2 (N-CoR2) (Silencing mediator of retinoic acid and thyroid hormone receptor) (SMRT) (SMRTe) (Thyroid-, retinoic-acid-receptor-associated corepressor) (T3 receptor- associating factor) (TRAC).

WNK4_MOUSE
θ value	3.0926 (rank : 19)	NC score	0.002019 (rank : 26)
Query Neighborhood Hits	26	Target Neighborhood Hits	961	Shared Neighborhood Hits	6
SwissProt Accessions	Q80UE6, Q4VAC1, Q80XB5, Q80XN2, Q8R0N0, Q8R340	Gene names	Wnk4, Prkwnk4
Domain Architecture
Description	Serine/threonine-protein kinase WNK4 (EC 2.7.11.1) (Protein kinase with no lysine 4) (Protein kinase, lysine-deficient 4).

IQEC2_HUMAN
θ value	4.03905 (rank : 20)	NC score	0.012231 (rank : 17)
Query Neighborhood Hits	26	Target Neighborhood Hits	403	Shared Neighborhood Hits	4
SwissProt Accessions	Q5JU85, O60275	Gene names	IQSEC2, KIAA0522
Domain Architecture	No domain graphic available
Description	IQ motif and Sec7 domain-containing protein 2.

IQEC2_MOUSE
θ value	4.03905 (rank : 21)	NC score	0.012307 (rank : 16)
Query Neighborhood Hits	26	Target Neighborhood Hits	417	Shared Neighborhood Hits	5
SwissProt Accessions	Q5DU25	Gene names	Iqsec2, Kiaa0522
Domain Architecture	No domain graphic available
Description	IQ motif and Sec7 domain-containing protein 2.

CD2AP_MOUSE
θ value	5.27518 (rank : 22)	NC score	0.006968 (rank : 21)
Query Neighborhood Hits	26	Target Neighborhood Hits	336	Shared Neighborhood Hits	4
SwissProt Accessions	Q9JLQ0, O88903, Q8K4Z1, Q8VCI9	Gene names	Cd2ap, Mets1
Domain Architecture
Description	CD2-associated protein (Mesenchyme-to-epithelium transition protein with SH3 domains 1) (METS-1).

AL2S8_MOUSE
θ value	6.88961 (rank : 23)	NC score	0.012016 (rank : 18)
Query Neighborhood Hits	26	Target Neighborhood Hits	31	Shared Neighborhood Hits	2
SwissProt Accessions	Q8VHI4	Gene names	Als2cr8, Carf
Domain Architecture	No domain graphic available
Description	Amyotrophic lateral sclerosis 2 chromosomal region candidate gene 8 protein homolog (Calcium-response factor) (CaRF).

MAGI2_MOUSE
θ value	6.88961 (rank : 24)	NC score	0.004563 (rank : 23)
Query Neighborhood Hits	26	Target Neighborhood Hits	370	Shared Neighborhood Hits	4
SwissProt Accessions	Q9WVQ1, Q3UH81, Q6GT88, Q8BYT1, Q8CA85	Gene names	Magi2, Acvrinp1, Aip1, Arip1
Domain Architecture
Description	Membrane-associated guanylate kinase, WW and PDZ domain-containing protein 2 (Membrane-associated guanylate kinase inverted 2) (MAGI-2) (Atrophin-1-interacting protein 1) (Activin receptor-interacting protein 1) (Acvrip1).

PCLO_HUMAN
θ value	8.99809 (rank : 25)	NC score	0.002526 (rank : 25)
Query Neighborhood Hits	26	Target Neighborhood Hits	2046	Shared Neighborhood Hits	13
SwissProt Accessions	Q9Y6V0, O43373, O60305, Q9BVC8, Q9UIV2, Q9Y6U9	Gene names	PCLO, ACZ, KIAA0559
Domain Architecture
Description	Protein piccolo (Aczonin).

TFEB_HUMAN
θ value	8.99809 (rank : 26)	NC score	0.006839 (rank : 22)
Query Neighborhood Hits	26	Target Neighborhood Hits	111	Shared Neighborhood Hits	2
SwissProt Accessions	P19484, Q709B3, Q7Z6P9, Q9BRJ5, Q9UJD8	Gene names	TFEB
Domain Architecture
Description	Transcription factor EB.

Hits sorted by NC score

VMD2_HUMAN
NC score	0.999962 (rank : 1)	θ value	0 (rank : 1)
Query Neighborhood Hits	26	Target Neighborhood Hits	26	Shared Neighborhood Hits	26
SwissProt Accessions	O76090, O75904, Q8IUR9, Q8IZ80	Gene names	VMD2
Domain Architecture
Description	Bestrophin-1 (Vitelliform macular dystrophy protein 2) (TU15B).

VMD2_MOUSE
NC score	0.992437 (rank : 2)	θ value	0 (rank : 2)
Query Neighborhood Hits	26	Target Neighborhood Hits	15	Shared Neighborhood Hits	7
SwissProt Accessions	O88870, Q6H1V0	Gene names	Vmd2, Bmd1
Domain Architecture	No domain graphic available
Description	Bestrophin-1 (Vitelliform macular dystrophy protein 2 homolog).

VMDL2_HUMAN
NC score	0.985886 (rank : 3)	θ value	2.69525e-129 (rank : 7)
Query Neighborhood Hits	26	Target Neighborhood Hits	8	Shared Neighborhood Hits	7
SwissProt Accessions	Q8NFU0	Gene names	VMD2L2
Domain Architecture
Description	Bestrophin-3 (Vitelliform macular dystrophy 2-like protein 2).

VMDL1_MOUSE
NC score	0.984085 (rank : 4)	θ value	2.35023e-141 (rank : 6)
Query Neighborhood Hits	26	Target Neighborhood Hits	19	Shared Neighborhood Hits	7
SwissProt Accessions	Q8BGM5, Q8VCM0	Gene names	Vmd2l1
Domain Architecture
Description	Bestrophin-2 (Vitelliform macular dystrophy 2-like protein 1).

VMDL1_HUMAN
NC score	0.983462 (rank : 5)	θ value	2.35023e-141 (rank : 5)
Query Neighborhood Hits	26	Target Neighborhood Hits	19	Shared Neighborhood Hits	7
SwissProt Accessions	Q8NFU1, Q9NXP0	Gene names	VMD2L1
Domain Architecture
Description	Bestrophin-2 (Vitelliform macular dystrophy 2-like protein 1).

VMDL3_MOUSE
NC score	0.978253 (rank : 6)	θ value	3.50381e-145 (rank : 4)
Query Neighborhood Hits	26	Target Neighborhood Hits	30	Shared Neighborhood Hits	8
SwissProt Accessions	Q6H1V1	Gene names	Vmd2l3
Domain Architecture	No domain graphic available
Description	Bestrophin-4 (Vitelliform macular dystrophy 2-like protein 3).

VMDL3_HUMAN
NC score	0.977047 (rank : 7)	θ value	3.50381e-145 (rank : 3)
Query Neighborhood Hits	26	Target Neighborhood Hits	32	Shared Neighborhood Hits	7
SwissProt Accessions	Q8N1M1, Q8N356, Q8NFT9, Q9BR80	Gene names	VMD2L3
Domain Architecture
Description	Bestrophin-4 (Vitelliform macular dystrophy 2-like protein 3).

ATE1_HUMAN
NC score	0.046737 (rank : 8)	θ value	0.0961366 (rank : 8)
Query Neighborhood Hits	26	Target Neighborhood Hits	11	Shared Neighborhood Hits	2
SwissProt Accessions	O95260, O95261, Q8WW04	Gene names	ATE1
Domain Architecture
Description	Arginyl-tRNA--protein transferase 1 (EC 2.3.2.8) (R-transferase 1) (Arginyltransferase 1) (Arginine-tRNA--protein transferase 1).

UBP10_HUMAN
NC score	0.026889 (rank : 9)	θ value	0.163984 (rank : 10)
Query Neighborhood Hits	26	Target Neighborhood Hits	153	Shared Neighborhood Hits	3
SwissProt Accessions	Q14694, Q9BWG7, Q9NSL7	Gene names	USP10, KIAA0190
Domain Architecture
Description	Ubiquitin carboxyl-terminal hydrolase 10 (EC 3.1.2.15) (Ubiquitin thioesterase 10) (Ubiquitin-specific-processing protease 10) (Deubiquitinating enzyme 10).

TULP4_MOUSE
NC score	0.020710 (rank : 10)	θ value	1.06291 (rank : 14)
Query Neighborhood Hits	26	Target Neighborhood Hits	247	Shared Neighborhood Hits	4
SwissProt Accessions	Q9JIL5, Q8CA75, Q922C2	Gene names	Tulp4, Tusp
Domain Architecture
Description	Tubby-like protein 4 (Tubby superfamily protein).

CYTSA_HUMAN
NC score	0.017808 (rank : 11)	θ value	0.0961366 (rank : 9)
Query Neighborhood Hits	26	Target Neighborhood Hits	1138	Shared Neighborhood Hits	5
SwissProt Accessions	Q69YQ0, O15081	Gene names	CYTSA, KIAA0376
Domain Architecture	No domain graphic available
Description	Cytospin-A (NY-REN-22 antigen).

IF4G1_HUMAN
NC score	0.015539 (rank : 12)	θ value	1.38821 (rank : 15)
Query Neighborhood Hits	26	Target Neighborhood Hits	347	Shared Neighborhood Hits	5
SwissProt Accessions	Q04637, O43177, O95066, Q5HYG0, Q6ZN21, Q8N102	Gene names	EIF4G1, EIF4G, EIF4GI
Domain Architecture
Description	Eukaryotic translation initiation factor 4 gamma 1 (eIF-4-gamma 1) (eIF-4G1) (eIF-4G 1) (p220).

CYTSA_MOUSE
NC score	0.015434 (rank : 13)	θ value	0.62314 (rank : 13)
Query Neighborhood Hits	26	Target Neighborhood Hits	1104	Shared Neighborhood Hits	3
SwissProt Accessions	Q2KN98, Q8CHF9	Gene names	Cytsa, Kiaa0376
Domain Architecture	No domain graphic available
Description	Cytospin-A.

EFS_MOUSE
NC score	0.014075 (rank : 14)	θ value	3.0926 (rank : 17)
Query Neighborhood Hits	26	Target Neighborhood Hits	389	Shared Neighborhood Hits	8
SwissProt Accessions	Q64355	Gene names	Efs, Sin
Domain Architecture
Description	Embryonal Fyn-associated substrate (SRC-interacting protein) (Signal- integrating protein).

PLXB1_MOUSE
NC score	0.013462 (rank : 15)	θ value	0.47712 (rank : 12)
Query Neighborhood Hits	26	Target Neighborhood Hits	259	Shared Neighborhood Hits	3
SwissProt Accessions	Q8CJH3, Q6ZQC3, Q80ZZ1	Gene names	Plxnb1, Kiaa0407
Domain Architecture	No domain graphic available
Description	Plexin-B1 precursor.

IQEC2_MOUSE
NC score	0.012307 (rank : 16)	θ value	4.03905 (rank : 21)
Query Neighborhood Hits	26	Target Neighborhood Hits	417	Shared Neighborhood Hits	5
SwissProt Accessions	Q5DU25	Gene names	Iqsec2, Kiaa0522
Domain Architecture	No domain graphic available
Description	IQ motif and Sec7 domain-containing protein 2.

IQEC2_HUMAN
NC score	0.012231 (rank : 17)	θ value	4.03905 (rank : 20)
Query Neighborhood Hits	26	Target Neighborhood Hits	403	Shared Neighborhood Hits	4
SwissProt Accessions	Q5JU85, O60275	Gene names	IQSEC2, KIAA0522
Domain Architecture	No domain graphic available
Description	IQ motif and Sec7 domain-containing protein 2.

AL2S8_MOUSE
NC score	0.012016 (rank : 18)	θ value	6.88961 (rank : 23)
Query Neighborhood Hits	26	Target Neighborhood Hits	31	Shared Neighborhood Hits	2
SwissProt Accessions	Q8VHI4	Gene names	Als2cr8, Carf
Domain Architecture	No domain graphic available
Description	Amyotrophic lateral sclerosis 2 chromosomal region candidate gene 8 protein homolog (Calcium-response factor) (CaRF).

NCOR2_MOUSE
NC score	0.010198 (rank : 19)	θ value	3.0926 (rank : 18)
Query Neighborhood Hits	26	Target Neighborhood Hits	790	Shared Neighborhood Hits	4
SwissProt Accessions	Q9WU42, Q9WU43, Q9WUC1	Gene names	Ncor2, Smrt
Domain Architecture
Description	Nuclear receptor corepressor 2 (N-CoR2) (Silencing mediator of retinoic acid and thyroid hormone receptor) (SMRT) (SMRTe) (Thyroid-, retinoic-acid-receptor-associated corepressor) (T3 receptor- associating factor) (TRAC).

CSPG2_MOUSE
NC score	0.007615 (rank : 20)	θ value	2.36792 (rank : 16)
Query Neighborhood Hits	26	Target Neighborhood Hits	711	Shared Neighborhood Hits	5
SwissProt Accessions	Q62059, Q62058, Q9CUU0	Gene names	Cspg2
Domain Architecture
Description	Versican core protein precursor (Large fibroblast proteoglycan) (Chondroitin sulfate proteoglycan core protein 2) (PG-M).

CD2AP_MOUSE
NC score	0.006968 (rank : 21)	θ value	5.27518 (rank : 22)
Query Neighborhood Hits	26	Target Neighborhood Hits	336	Shared Neighborhood Hits	4
SwissProt Accessions	Q9JLQ0, O88903, Q8K4Z1, Q8VCI9	Gene names	Cd2ap, Mets1
Domain Architecture
Description	CD2-associated protein (Mesenchyme-to-epithelium transition protein with SH3 domains 1) (METS-1).

TFEB_HUMAN
NC score	0.006839 (rank : 22)	θ value	8.99809 (rank : 26)
Query Neighborhood Hits	26	Target Neighborhood Hits	111	Shared Neighborhood Hits	2
SwissProt Accessions	P19484, Q709B3, Q7Z6P9, Q9BRJ5, Q9UJD8	Gene names	TFEB
Domain Architecture
Description	Transcription factor EB.

MAGI2_MOUSE
NC score	0.004563 (rank : 23)	θ value	6.88961 (rank : 24)
Query Neighborhood Hits	26	Target Neighborhood Hits	370	Shared Neighborhood Hits	4
SwissProt Accessions	Q9WVQ1, Q3UH81, Q6GT88, Q8BYT1, Q8CA85	Gene names	Magi2, Acvrinp1, Aip1, Arip1
Domain Architecture
Description	Membrane-associated guanylate kinase, WW and PDZ domain-containing protein 2 (Membrane-associated guanylate kinase inverted 2) (MAGI-2) (Atrophin-1-interacting protein 1) (Activin receptor-interacting protein 1) (Acvrip1).

SPEG_HUMAN
NC score	0.004118 (rank : 24)	θ value	0.365318 (rank : 11)
Query Neighborhood Hits	26	Target Neighborhood Hits	1762	Shared Neighborhood Hits	7
SwissProt Accessions	Q15772, Q27J74, Q695L1, Q6FGA6, Q6ZQW1, Q6ZTL8, Q9P2P9	Gene names	SPEG, APEG1, KIAA1297
Domain Architecture
Description	Striated muscle preferentially expressed protein kinase (EC 2.7.11.1) (Aortic preferentially expressed protein 1) (APEG-1).

PCLO_HUMAN
NC score	0.002526 (rank : 25)	θ value	8.99809 (rank : 25)
Query Neighborhood Hits	26	Target Neighborhood Hits	2046	Shared Neighborhood Hits	13
SwissProt Accessions	Q9Y6V0, O43373, O60305, Q9BVC8, Q9UIV2, Q9Y6U9	Gene names	PCLO, ACZ, KIAA0559
Domain Architecture
Description	Protein piccolo (Aczonin).

WNK4_MOUSE
NC score	0.002019 (rank : 26)	θ value	3.0926 (rank : 19)
Query Neighborhood Hits	26	Target Neighborhood Hits	961	Shared Neighborhood Hits	6
SwissProt Accessions	Q80UE6, Q4VAC1, Q80XB5, Q80XN2, Q8R0N0, Q8R340	Gene names	Wnk4, Prkwnk4
Domain Architecture
Description	Serine/threonine-protein kinase WNK4 (EC 2.7.11.1) (Protein kinase with no lysine 4) (Protein kinase, lysine-deficient 4).